Graduate Bulletin

Spring 2022

Doctoral Program in Anthropological Sciences

Distinguished Professors

Fleagle, John G.,1 Distinguished Professor, Ph.D., 1976, Harvard University: Primate and human evolution; primate behavior and ecology; functional morphology; growth and development.

Grine, Frederick E.,2 Distinguished Professor, Ph.D., 1984, University of the Witwatersrand, South Africa: Hominid evolution; functional morphology of the masticatory apparatus; diet reconstruction; dental anthropology; mineralized tissues.

Wright, Patricia C.,2 Distinguished Service Professor, Ph.D., 1985, City University of New York: Primate behavior and ecology; rainforest conservation; Madagascar.


Davalos, Liliana M.,3 Ph.D., 2004, Columbia University: Phylogenetics, tropical deforestation.

Hicks, David,2 D.Phil., 1973, Oxford University, Great Britain: Politics; ritual; literature; East Timor; Insular Southeast Asia.

Koenig, Andreas,2 Ph.D., 1992, Georg-August University, Göttingen, Germany: Primate behavioral ecology; social evolution; reproductive strategies; Asia.

Lynch, Heather, Ph.D. 2006, Harvard University: Quantitative ecology, geospatial modeling, remote sensing and GIS

Martin, Lawrence B.,4 Director of TBI, Ph.D., 1983, University of London, Great Britain: Hominoid evolution; enamel thickness; enamel microstructure and development.

Shea, John J., Ph.D.,2 1991, Harvard University: Paleoanthropology, archaeology; lithic analysis; Southwest Asia; Eastern Africa; human dispersals.

Associate Professors

Harmand, Sonia.,2,4 Ph.D., 2005, Paris X, France: Early stone age archaeology; lithic technology; cognition; primate archaeology; Africa.

Hildebrand, Elisabeth,2,4 Ph.D., 2003, Washington University: Origins of agriculture; paleoethnobotany; ethnoarchaeology; Africa.

Markham, A. Catherine,2 Ph.D., 2012, Princeton University: Behavioral ecology; maternal care; spatial ecology; wild primates.

Newman, Elizabeth Terese,5 Ph.D., 2008, Yale University: Zooarchaeology; historical archaeology; household archaeology; gender; Mexico.

Rasbury, Troy, E.,6 Ph.D., 1998, Stony Brook University: Sedimentary geochemistry; chronostratigraphy; geochronology.

Rossie, James B.,2 Ph.D., 2003, Yale University: Primate evolution; miocene hominoids; cranial anatomy; East Africa.

Ruf, Gregory A.,7 Ph.D., 1994, Columbia University: Social organization; politics and history; cultural ecology; ethnographic methods and writing; China.

Seiffert, Erik R.,1 Ph.D., 2003, Duke University: Evolution of primates and other placental mammals.

Smaers, Jeroen B.,2 Ph.D., 2010, University of Cambridge, England: Brain evolution, phylogenetic comparative methodology, macroevolutionary morphology.

Turner, Alan H.,1 Ph.D., 2008, Columbia University: Phylogenetic methods, dinosaur and crocodylomorph evolution, morphological data, paleontology

Twiss, Katheryn C.,2 Ph.D., 2003, University of California, Berkeley: Archaeology of food; zooarchaeology; origins of agriculture and social complexity; Southwest Asia.

Veeramah, Krishna R.,3 Ph.D., 2008, University College London, Great Britain: primate evolution; genomics; population genetics; sub-Saharan Africa, Ancient DNA.

Research Professor

Leakey Meave G.,4 PhD., 1968, University of North Wales: Primate evolution; palaeoecology and evolution of African mammals.

Research Associate Professor

Borries, Carola,2 Ph.D., 1989, Georg-August University, Göttingen, Germany: Socio-ecology; reproduction; life history; Asian primates.

Assistant Professors

Frouin, Marine,6 Ph.D., 2014, Université Bordeaux Montaigne, France: Luminescence dating, geochronology, chronostratigraphy

Henkes, Gregory A.,6 Ph.D., 2014, Johns Hopkins University: Stable isotope geochemistry, paleoclimatology, biogeochemistry.

Lu, Amy,2 Ph.D., 2009, Stony Brook University: Behavioral endocrinology, socioecology, sexual selection, growth and development, life history

Maiolino, Stephanie A.,1 Ph.D., 2015, Stony Brook University: primate evolution, comparative morphology, hands and feet, locomotion

Percival, Christopher J.,2 Ph.D., 2013, Penn State University: Skull development and evolution, craniofacial disease, geometric morphometrics, computed tomography image analysis

Russo, Gabrielle A.,2 Ph.D., 2013, University of Texas at Austin: Functional morphology of the axial skeleton, primate and human evolution, locomotion, ontogeny.

Smiley, Tara M.,3 Ph.D., 2016, University of Michigan: Diversity dynamics and paleoecology of fossil mammals, biogeography, stable isotope ecology.

Vitek, Natasha S.,3 Ph.D., 2019, University of Florida: Evolution and intraspecific variation in vertebrate fossils, primarily mammals and turtles.

Yager, Karina A.,8 Ph.D., 2009, Yale University: Pastoralism; cultural ecology; remote sensing; climate change and society; Andean studies.

Research Assistant Professors

Lewis, Jason,2,4 Ph.D., 2011, Stanford University: Paleoecology, paleontology, human evolution, zooarchaeology, Old World Paleolithic archaeology.

Nengo, Isaiah, Ph.D., 1993, Harvard University: Primate paleontology; ape and human  evolution

Emeriti Faculty

Demes, Brigitte,1 Professor Emerita, Ph.D., 1982, Ruhr University, Bochum, Germany: Biomechanics; functional morphology; allometry; primates.

Jungers, William L.,1 Distinguished Teaching Professor Emeritus, Ph.D., 1976, University of Michigan: Primate and human evolution; functional morphology; biomechanics.

Stone, Elizabeth C.,2 Ph.D., 1979, University of Chicago: Old World archaeology; state formation; ancient economy and society; remote sensing and GIS; Near East.

Zimansky, Paul E.,5 Ph.D., 1980, University of Chicago: History and archaeology of the Near East; ancient imperialism; Urartian, Anatolian and Mesopotamian civilizations.


Number of teaching assistants/graduate assistants/research assistants, fall 2019: 16 Number of graduate fellows, fall 2019: 13


  1. Department of Anatomical Sciences
  2. Department of Anthropology
  3. Department of Ecology and Evolution
  4. Turkana Basin Institute
  5. Department of History
  6. Department of Geosciences
  7. Department of Asian and Asian American Studies
  8. Sustainability Studies Program