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2021 Publications from the Department of Ecology and Evolution

  • Molly K Grace, Elizabeth L Bennett, H Reşit Akçakaya, Craig Hilton-Taylor, Michael Hoffmann, Richard Jenkins, EJ Milner-Gulland, Ana Nieto, Richard P Young, Barney Long, 2021, IUCN launches Green Status of Species: a new standard for species recovery, Oryx, Volume 55, Issue 5, pp.651-652, DOI: https://doi.org/10.1017/S0030605321000880.
  • Molly K Grace, H Resit Akçakaya, Elizabeth L Bennett, Thomas M Brooks, Anna Heath, Simon Hedges, Craig Hilton‐Taylor, Michael Hoffmann, Axel Hochkirch, Richard Jenkins, David A Keith, Barney Long, David P Mallon, Erik Meijaard, EJ Milner‐Gulland, Jon Paul Rodriguez, PJ Stephenson, Simon N Stuart, Richard P Young, Pablo Acebes, Joanna Alfaro‐Shigueto, Silvia Alvarez‐Clare, Raphali Rodlis Andriantsimanarilafy, Marina Arbetman, Claudio Azat, Gianluigi Bacchetta, Ruchi Badola, Luís MD Barcelos, Joao Pedro Barreiros, Sayanti Basak, Danielle J Berger, Sabuj Bhattacharyya, Gilad Bino, Paulo AV Borges, Raoul K Boughton, H Jane Brockmann, Hannah L Buckley, Ian J Burfield, James Burton, Teresa Camacho‐Badani, Luis Santiago Cano‐Alonso, Ruth H Carmichael, Christina Carrero, John P Carroll, Giorgos Catsadorakis, David G Chapple, Guillaume Chapron, Gawsia Wahidunnessa Chowdhury, Louw Claassens, Donatella Cogoni, Rochelle Constantine, Christie Anne Craig, Andrew A Cunningham, Nishma Dahal, Jennifer C Daltry, Goura Chandra Das, Niladri Dasgupta, Alexandra Davey, Katharine Davies, Pedro Develey, Vanitha Elangovan, David Fairclough, Mirko Di Febbraro, Giuseppe Fenu, Fernando Moreira Fernandes, Eduardo Pinheiro Fernandez, Brittany Finucci, Rita Földesi, Catherine M Foley, Matthew Ford, Michael RJ Forstner, Néstor García, Ricardo Garcia‐Sandoval, Penny C Gardner, Roberto Garibay‐Orijel, Marites Gatan‐Balbas, Irene Gauto, Mirza Ghazanfar Ullah Ghazi, Stephanie S Godfrey, Matthew Gollock, Benito A González, Tandora D Grant, Thomas Gray, Andrew J Gregory, Roy HA van Grunsven, Marieka Gryzenhout, Noelle C Guernsey, Garima Gupta, Christina Hagen, Christian A Hagen, Madison B Hall, Eric Hallerman, Kelly Hare, Tom Hart, Ruston Hartdegen, Yvette Harvey‐Brown, Richard Hatfield, Tahneal Hawke, Claudia Hermes, Rod Hitchmough, Pablo Melo Hoffmann, Charlie Howarth, Michael A Hudson, Syed Ainul Hussain, Charlie Huveneers, Hélène Jacques, Dennis Jorgensen, Suyash Katdare, Lydia KD Katsis, Rahul Kaul, Boaz Kaunda‐Arara, Lucy Keith‐Diagne, Daniel T Kraus, Thales Moreira de Lima, Ken Lindeman, Jean Linsky, Edward Louis Jr, Anna Loy, Eimear Nic Lughadha, Jeffrey C Mangel, Paul E Marinari, Gabriel M Martin, Gustavo Martinelli, Philip JK McGowan, Alistair McInnes, Eduardo Teles Barbosa Mendes, Michael J Millard, Claire Mirande, Daniel Money, Joanne M Monks, Carolina Laura Morales, Nazia Naoreen Mumu, Raquel Negrao, Anh Ha Nguyen, Md Nazmul Hasan Niloy, Grant Leslie Norbury, Cale Nordmeyer, Darren Norris, Mark O'Brien, Gabriela Akemi Oda, Simone Orsenigo, Mark Evan Outerbridge, Stesha Pasachnik, Juan Carlos Pérez‐Jiménez, Charlotte Pike, Fred Pilkington, Glenn Plumb, Rita de Cassia Quitete Portela, Ana Prohaska, Manuel G Quintana, 2021, Testing a global standard for quantifying species recovery and assessing conservation impact, Conservation Biology,   https://doi.org/10.1111/cobi.13756.
  • H Reşit Akçakaya, Axel Hochkirch, Jason T Bried, Roy HA van Grunsven, John P Simaika, Geert De Knijf, Sérgio Henriques, 2021, Calculating population reductions of invertebrate species for IUCN Red List assessments, Journal of Insect Conservation, volume 25, pp.377–382.
  • Bilgecan Şen, H Reşit Akçakaya, 2021, Inter-specific Variability in Demographic Processes Affects Abundance-Occupancy Relationships, bioRxiv, doi: https://doi.org/10.1101/2020.09.03.282004.
  • Damien A Fordham, Stuart C Brown, H Resit Akcakaya, Barry W Brook, Sean Haythorne, Andrea Manica, Kevin T Shoemaker, Jeremy J Austin, Benjamin Blonder, Julia Pilowsky, Carsten Rahbek, David Nogues-Bravo, 2021, Humans hastened the range collapse and extinction of woolly mammoth, bioRxiv, doi: https://doi.org/10.1101/2021.02.17.431706.
  • Molly K.Grace, H. Resit Akçakaya, Joseph W.Bull, Christina Carrero, Katharine Davies, Simon Hedges, Michael Hoffmann, Barney Long, Eimear M. Nic Lughadha, Gabriel M.Martin, Fred Pilkington, Malin C.Rivers, Richard P.Young, E.J.Milner-Gulland, 2021, Building robust, practicable counterfactuals and scenarios to evaluate the impact of species conservation interventions using inferential approaches, Biological Conservation, Volume 261, https://doi.org/10.1016/j.biocon.2021.109259.
  • Emily M Herstoff, Cédric L Meunier, Maarten Boersma, Stephen B Baines, 2021, Leveraging differences in multiple prey traits allows selective copepods to meet their threshold elemental ratios, Limnology and Oceanography, https://doi.org/10.1002/lno.11800.
  • Rafael D’Andrea, James P O’Dwyer, 2021, Competition for space in a structured landscape: The effect of seed limitation on coexistence under a tolerance‐fecundity trade‐off, Journal of Ecology, https://doi.org/10.1111/1365-2745.13613.
  • Armenteras-Pascual, D., L.M. Dávalos, J.S. Barreto, A. Miranda, A. Hernández-Moreno, C. Zamorano-Elgueta, T. M. González-Delgado, M. C. Meza, Javier Retana-Alumbreros. Fire-induced loss of the world’s most biodiverse forests in Latin America. Science Advances 7(33), eabd3357 DOI: 10.1126/sciadv.abd3357.
  • Santana, S.E.*, Z.A. Kaliszewska*, L.B. Leiser-Miller, M.E. Lauterbur, J.H. Arbour, L.M. Dávalos, J.A. Riffell. Fruit odorants mediate co-specialization in a multispecies plant-animal mutualism. Proceedings of the Royal Society B: Biological Sciences 288: 28820210312 DOI: 10.1098/rspb.2021.0312. *Equal contribution.
  • Lertxundi, U., S. Domingo-Echaburu, L.M. Dávalos, G. Orive. Drug pollution & Sustainable Developing Goals. Science of the Total Environment.
  • Yohe, L.R., L.B. Leiser-Miller, Z.A. Kaliszewska, P. Donat, S.E. Santana, L.M. Dávalos. Diversity in olfactory receptor repertoires is associated with dietary specialization in a genus of frugivorous bat. G3: Genes|Genomes|Genetics. DOI: 10.1093/g3journal/jkab260.
  • Moreno Santillan, D., T. Lama, Y.G. Guerrero, A. Brown, P. Donat, H. Zhao, S. Rossiter, L.R. Yohe, J. Potter, E.C. Teeling, S. Vernes, K.T.J. Davies, E. Myers, G. Hughes, Z. Huang, F. G. Hoffmann, A.P. Corthals, D. Ray*, L.M. Dávalos*. Large-Scale Genome Sampling Reveals Unique Immunity and Metabolic Adaptations in Bats. Molecular Ecology. DOI: 10.1111/MEC.16027. *Corresponding authors.
  • Hall, R.P., G. Mutumi, B.P. Hedrick, L.R. Yohe, A. Sadier, K.T.J. Davies, S.J. Rossiter, K. Sears, L.M. Dávalos, E.R. Dumont. Find the Food First: An Omnivorous Sensory Morphotype Predates Biomechanical Specialization for Plant Based Diets in Phyllostomid Bats. Evolution. DOI: 10.1111/evo.14270.
  • Murillo-Sandoval, P., E. Gjerdseth, C. Correa-Ayram, D. Wrathall, J. Van Den Hoek, L. M. Dávalos, R. Kennedy. No peace for the forest: Rapid, widespread land changes in the Andes-Amazon region following the Colombian civil war. Global Environmental Change 69: 102283. DOI: 10.1016/j.gloenvcha.2021.102283.
  • Turvey, S.T.*, C. Duncan, N.S. Upham, X. Harrison, L.M. Dávalos*. Where the wild things were: intrinsic and extrinsic extinction predictors in the world’s most depleted mammal fauna. Proceedings of the Royal Society B: Biological Sciences 288: 20202905. DOI: 10.1098/rspb.2020.2905. *Corresponding authors.
  • Lim, M., K. Bi, C. Witt, C. Graham, L. Dávalos. Pervasive genomic signatures of local adaptation to altitude across highland specialist Andean hummingbird populations. Journal of Heredity. DOI: 10.1093/jhered/esab008.
  • Potter, J.H.T., K.T J. Davies, L.R. Yohe, K. Warren, G. Tsagkogeorga, B.K. Lim, M. dos Reis, L.M. Dávalos, S.J. Rossite*. Dietary diversification and specialisation in neotropical bats facilitated by early molecular evolution. Molecular Biology and Evolution. DOI: 10.1093/molbev/msab028.
  • Potter, J.H.T.*, R. Drinkwater, K.T J. Davies, M.C.W. Lim, L.R. Yohe, X. Zhang, H. Chi, I. Levantis, B.K. Lim, C.C. Witt, G. Tsagkogeorga, M. dos Reis, Y. Liu, W. Furey, M.J. Whitley, D. Aksentijevic, L.M. Dávalos, S.J. Rossiter*, 2021, Nectar-feeding bats and birds show parallel molecular adaptations in sugar metabolism enzymes, Current Biology, DOI: 10.1016/j.cub.2021.08.018, *Corresponding authors.
  • Daniel E Dykhuizen, 2021, Public Health and Paul De Kruif, The Quarterly Review of Biology, 96, pp.127-132.
  • O’Dea, R.E., Lagisz, M., Jennions, M.D., Koricheva, J., Noble, D.W.A., Parker, T.H., Gurevitch, J., Page, M.J., Stewart, G., Moher, D., Nakagawa, S., n.d. Preferred reporting items for systematic reviews and meta-analyses in ecology and evolutionary biology: a PRISMA extension. Biol. Rev. 2021. https://doi.org/10.1111/brv.12721.
  • Phoebe L. Zarnetske*, Jessica Gurevitch* [lead authors], Janet Franklin, Peter Groffman, Cheryl Harrison, Jessica Hellmann, Forrest M. Hoffman, Shan Kothari, Alan Robock, Simone Tilmes, Daniele Visioni, Jin Wu, Lili Xia, Cheng‐En Yang. 2021. Potential ecological impacts of climate intervention by reflecting sunlight to cool Earth. Proceedings of the National Academy of Sciences 2021 Vol. 118 No. 15 e1921854118;   https://doi.org/10.1073/pnas.1921854118.
  • Gurevitch, J., Jadotte, Y. and A. Moyer. 2021. Meta-analysis in the Social Sciences. (Chapter) The Cambridge Handbook of Research Methods and Statistics for the Social and Behavioral Sciences , Volume 1 edited by Austin Lee Nichols; John Edlund. In press.
  • Tonjes, David and Jessica Gurevitch. 21 May 2021. Trash Talk: Garbage and the Infrastructure Bill (retitled, Better ways to manage LI’s trash). Newsday Opinion article.
  • Riley C Pena, Jeffrey Levinton, 2021, Two species of fiddler crab show no cost of bearing a sexually selected giant claw in predator escape behaviors, Journal of Experimental Marine Biology and Ecology, Volume 541, August 2021, 151570, https://doi.org/10.1016/j.jembe.2021.151570.
  • Thomas S Bianchi, Robert C Aller, Trisha B Atwood, Craig J Brown, Luis A Buatois, Lisa A Levin, Jeffrey S Levinton, Jack J Middelburg, Elise S Morrison, Pierre Regnier, Michael R Shields, Paul VR Snelgrove, Erik E Sotka, Ryan RE Stanley, 2021, What global biogeochemical consequences will marine animal–sediment interactions have during climate change? Elementa: Science of the Anthropocene (2021) 9 (1): 00180, https://doi.org/10.1525/elementa.2020.00180.
  • Jeffrey S Levinton, Brooke Arena, 2021, Direct and biometrical estimates of the closing force of major claws of the sand fiddler crab Leptuca pugilator (Bosc, 1801) (Decapoda: Brachyura: Ocypodidae): Support for the weakening combatant hypothesis, Journal of Crustacean Biology, Volume 41, Issue 2, June 2021, ruab007, https://doi.org/10.1093/jcbiol/ruab007
  • Tom Hart, Fiona Jones, Caitlin Black, Chris Lintott, Casey Youngflesh, Heather Lynch, Alasdair Davies, Eamonn Maguire, Andrew Zisserman, Carlos Arteta, Peter Barham, Colin Southwell, Louise Emmerson, Mark Jessopp, 2021, Time-lapse imagery is cheap and timely in the fight against colonial species' decline, Authorea Preprints, DOI: 10.22541/au.162686380.02020424/v1.
  • LR Pertierra, F Santos-Martin, KA Hughes, C Avila, JO Caceres, D De Filippo, S Gonzalez, SM Grant, H Lynch, C Marina-Montes, A Quesada, P Tejedo, T Tin, J Benayas, 2021, Ecosystem services in Antarctica: Global assessment of the current state, future challenges and managing opportunities, Ecosystem Services, Vol.49, pp.101299, https://doi.org/10.1016/j.ecoser.2021.101299.
  • Alex Borowicz, Heather J Lynch, Tyler Estro, Catherine Foley, Bento Gonçalves, Katelyn B Herman, Stephanie K Adamczak, Ian Stirling, Lesley Thorne, 2021, Social sensors for wildlife: Ecological opportunities in the era of camera ubiquity, Frontiers in Marine Science, Vol.8, pp.385, https://doi.org/10.3389/fmars.2021.645288.
  • Aymen Al-Saadi, Ioannis Paraskevakos, Bento Collares Gonçalves, Heather J Lynch, Shantenu Jha, Matteo Turilli, 2021, Comparing workflow application designs for high resolution satellite image analysis, Future Generation Computer Systems, https://doi.org/10.1016/j.future.2021.04.023.
  • Michael Schrimpf, Heather Lynch, 2021, The role of wind fetch in structuring Antarctic seabird breeding occupancy, Ibis, Vol.163, No.2, pp.695-705, https://doi.org/10.1111/ibi.12910.
  • Casey Youngflesh, Fiona M Jones, Heather J Lynch, Joan Arthur, Zuzana Ročkaiová, Holly R Torsey, Tom Hart, 2021, Large‐scale assessment of intra‐and inter‐annual breeding success using a remote camera network, Remote Sensing in Ecology and Conservation, Vol.7, No.1, pp.97-108, https://doi.org/10.1002/rse2.171.
  • Jonathon Handley, Marie-Morgane Rouyer, Elizabeth J Pearmain, Victoria Warwick-Evans, Katharina Teschke, Jefferson Hinke, Heather Lynch, Louise Emmerson, Colin Southwell, Gary Griffith, Cesar A Cardenas, Aldina Franco, Philip N Trathan, Maria P Dias, 2021, Marine important bird and biodiversity areas for penguins in Antarctica: Targets for conservation action, Frontiers in Marine Science, Vol.7, pp.1190, https://doi.org/10.3389/fmars.2020.602972.
  • Bento C Gonçalves, Heather J Lynch, 2021, Fine-Scale Sea Ice Segmentation for High-Resolution Satellite Imagery with Weakly-Supervised CNNs, Remote Sensing, 2021, 13(18), 3562; https://doi.org/10.3390/rs13183562.
  • Noah Strycker, Alex Borowicz, Michael Wethington, Steven Forrest, Vikrant Shah, Yang Liu, Hanumant Singh, Heather J Lynch, 2021, Fifty-year change in penguin abundance on Elephant Island, South Shetland Islands, Antarctica: results of the 2019–20 census, Polar Biology, 44, pages45–56 (2021).
  • Roberto Bertolini, Stephen J Finch, Ross H Nehm, 2021, Enhancing data pipelines for forecasting student performance: integrating feature selection with cross-validation, International Journal of Educational Technology in Higher Education, volume 18, pp.1-23, Article number: 44 (2021).
  • Karen Nicole Pelletreau, Tessa Andrews, Norris Armstrong, Mary A Bedell, Farahad Dastoor, Neta Dean, Susan Erster, Cori Fata-Hartley, Nancy Guild, Hamish Greig, David Hall, Jennifer K Knight, Donna Koslowsky, Paula Lemons, Jennifer Martin, Jill McCourt, John Merrill, Rosa Moscarella, Ross Nehm, Robert Northington, Brian Olsen, Luanna Prevost, Jon Stolzfus, Mark Urban-Lurain, Michelle K Smith, 2021, A clicker-based case study that untangles student thinking about the processes in the central dogma, CourseSource.
  • Xiaoming Zhai, Lehong Shi, Ross H Nehm, 2021, A meta-analysis of machine learning-based science assessments: factors impacting machine-human score agreements, Journal of Science Education and Technology, volume 30, pages361–379 (2021).
  • Roberto Bertolini, Stephen J Finch, Ross H Nehm, 2021, Testing the impact of novel assessment sources and machine learning methods on predictive outcome modeling in undergraduate biology, Journal of Science Education and Technology, volume 30, pages193–209 (2021).
  • Gena C Sbeglia, Justin A Goodridge, Lucy H Gordon, Ross H Nehm, 2021, Are Faculty Changing? How Reform Frameworks, Sampling Intensities, and Instrument Measures Impact Inferences about Student-Centered Teaching Practices, CBE—Life Sciences Education, Vol.20, No.3 pp.ar39, https://doi.org/10.1187/cbe.20-11-0259.
  • Alexandra P Hooks, Dianna K Padilla, 2021, Introduced predator elicits population-specific responses from prey, Biological Invasions, 23, pp.477–490.
  • Niamh B O’Hara, Steven J Franks, Nolan C Kane, Silas Tittes, Joshua S Rest, 2021, Evolution of pathogen response genes associated with increased disease susceptibility during adaptation to an extreme drought in a Brassica rapa plant population, BMC Ecology and Evolution, 21, pp.1-11.
  • Joanne Figueiredo, Joshua Rest, Sarah Adamo, Rebecca Grella, James Dilger, 2021, Nicotine Stimulates Peristalsis in N. Vectensis: From Behavior to Nicotinic Receptor Genes, Research Square, DOI:https://doi.org/10.21203/rs.3.rs-195699/v1.
  • Elena Hamann, Christopher S Pauli, Zoé Joly‐Lopez, Simon C Groen, Joshua S Rest, Nolan C Kane, Michael D Purugganan, Steven J Franks, 2021, Rapid evolutionary changes in gene expression in response to climate fluctuations, Molecular ecology, 30, pp.193.206, https://doi.org/10.1111/mec.15583.
  • Jeroen B Smaers, Ryan S Rothman, Daphne R Hudson, Amy M Balanoff, Brian Beatty, Dina KN Dechmann, Dorien de Vries, Jacob C Dunn, John G Fleagle, Christopher C Gilbert, Anjali Goswami, Andrew N Iwaniuk, William L Jungers, Max Kerney, Daniel T Ksepka, Paul R Manger, CS Mongle, F James Rohlf, N Adam Smith, Christophe Soligo, Vera Weisbecker, Kamran Safi, 2021, The evolution of mammalian brain size, SCIENCE ADVANCES, 28 Apr 2021, Vol 7, Issue 18, DOI: 10.1126/sciadv.abe2101.
  • F James Rohlf, 2021, Sampling Theory: For the Ecological and Natural Resource Sciences, QUARTERLY REVIEW OF BIOLOGY, Vo.96, No.1, pp.45-45.
  • F James Rohlf, 2021, Why Clusters and Other Patterns Can Seem to be Found in Analyses of High-Dimensional Data, Evolutionary Biology, volume 48, pages1–16 (2021).
  • T Akther, CG Easson, R Collin, RW Thacker, 2021, Comparing adult and larval microbiomes in the tropical sponges Neopetrosia sigmafera and Xestospongia bocatorensis, INTEGRATIVE AND COMPARATIVE BIOLOGY, Vol.61, pp.E1037-E1037.
  • Christopher J Freeman, Cole G Easson, Cara L Fiore, Robert W Thacker, 2021, Sponge–Microbe Interactions on Coral Reefs: Multiple Evolutionary Solutions to a Complex Environment, Frontiers in Marine Science, 20 July 2021, pp.917, https://doi.org/10.3389/fmars.2021.705053.
  • Joseph B Kelly, David E Carlson, Jun Siong Low, Tyler Rice, Robert W Thacker, 2021, The relationship between microbiomes and selective regimes in the sponge genus Ircinia, Frontiers in microbiology, Vol.12: 607289. doi: 10.3389/fmicb.2021.607289.
  • Matthew Houser, Abigail Sullivan, Tara Smiley, Ranjan Muthukrishnan, Elizabeth Grennan Browning, Adam Fudickar, Pascal Title, Jason Bertram, Maria Whiteman, 2021, What fosters the success of a transdisciplinary environmental research institute? Reflections from an interdisciplinary research cohort, Elementa: Science of the Anthropocene (2021) 9 (1): 00132. https://doi.org/10.1525/elementa.2020.00132.
  • Eric S Haag, John R True, 2021, Developmental System Drift, Evolutionary developmental biology: a reference guide, pp.99-110.
  • Garrett A Roberts Kingman, Deven N Vyas, Felicity C Jones, Shannon D Brady, Heidi I Chen, Kerry Reid, Mark Milhaven, Thomas S Bertino, Windsor E Aguirre, David C Heins, Frank A von Hippel, Peter J Park, Melanie Kirch, Devin M Absher, Richard M Myers, Federica Di Palma, Michael A Bell, David M Kingsley, Krishna R Veeramah, 2021, Predicting future from past: The genomic basis of recurrent and rapid stickleback evolution, SCIENCE ADVANCES, 18 Jun 2021, Vol 7, Issue 25, DOI: 10.1126/sciadv.abg5285.
  • A Žegarac, L Winkelbach, J Blöcher, Y Diekmann, M Krečković Gavrilović, M Porčić, B Stojković, L Milašinović, M Schreiber, D Wegmann, KR Veeramah, S Stefanović, J Burger, 2021, Ancient genomes provide insights into family structure and the heredity of social status in the early Bronze Age of southeastern Europe, Scientific Reports, volume 11, Article number: 10072 (2021).
  • Kerry Reid, Michael A Bell, Krishna R Veeramah, 2021, Threespine Stickleback: A Model System For Evolutionary Genomics, Annual Review of Genomics and Human Genetics, Vol. 22:357-383, https://doi.org/10.1146/annurev-genom-111720-081402.
  • Kyunghun Min, Thomas F Jannace, Haoyu Si, Krishna R Veeramah, John D Haley, James B Konopka, 2021, Integrative multi-omics profiling reveals cAMP-independent mechanisms regulating hyphal morphogenesis in Candida albicans, doi: https://doi.org/10.1101/2021.03.08.433668.
  • Walter G Joyce, Jérémy Anquetin, Edwin-Alberto Cadena, Julien Claude, Igor G Danilov, Serjoscha W Evers, Gabriel S Ferreira, Andrew D Gentry, Georgios L Georgalis, Tyler R Lyson, Adán Pérez-García, Márton Rabi, Juliana Sterli, Natasha S Vitek, James F Parham, 2021, A nomenclature for fossil and living turtles using phylogenetically defined clade names, Swiss Journal of Palaeontology, volume 140, Article number: 5 (2021).
  • Natasha S Vitek, Paul E Morse, Doug M Boyer, Suzanne G Strait, Jonathan I Bloch, 2021, Evaluating the responses of three closely related small mammal lineages to climate change across the Paleocene–Eocene thermal maximum, Paleobiology, Volume 47, Issue 3, August 2021, pp.464-486, DOI: https://doi.org/10.1017/pab.2021.12.